New Associations of Steppe Vegetation in National Park «Khvalynsky»
The article considers the issue of studying the steppe vegetation of the Khvalynsky National Park from the standpoint of ecologophytocenotic and ecologo-floristic approaches to the classification of vegetation. New associations Alysso tortuosi-Artemisietum salsoloidis Lysenko ass. nov. hoc loco, Euphorbio glareosae-Festucetum valesiacae ass. nov. hoc loco, Artemisio marschallianae-Stipetum pennatae Lysenko ass. nov. hoc loco, Astragalo varii-Bromopsietum inermis Lysenko ass. nov. hoc loco, Astragalo albicaulis-Stipetum lessingianae Lysenko ass. nov. hoc loco, Veronico prostrataeArtemisietum nitrosae Lysenko ass. nov. hoc loco and Galatello angustissimae-Spiraetum litwinowii Lysenko ass. nov. hoc loco are selected on the basis of the use of ecologo-floristic approach. Their diagnostic types, nomenclature types are given, the composition and structure, ecology and distribution of communities are characterized. Installed syntaxa are ranged to class Festuco-Brometea Br.-Bl. et Tx. ex Soo 1947, orders Festucetalia valesiacae Soo 1947, Helictotricho-Stipetalia Toman 1969 and Tanaceto achilleifoliiStipetalia lessingianae Lysenko et Mucina in Mucina 2016, unions Festucion valesiacae Klika 1931, Centaurion sumensis Golub et Uzhametskaya 2016 and Tanaceto achilleifolii-Stipion lessingianae Lysenko et Mucina in Mucina 2016.
1. Tarasov A. O. The structure of the vegetation cover of the Lower Volga region. Bull. MOIP Separate Biol., 1991, vol. 96, iss. 6, pp. 92–100 (in Russian).
2. Lavrenko E. M. Stepi [Steppes]. In: Rastitel'nost' evropeyskoy chasti SSSR [Vegetation of the European Part of the USSR]. Leningrad, Nauka Publ., Leningr. otd-nie, 1980, pp. 203–272 (in Russian).
3. Malysheva G. S., Malakhovskiy P. D. The structure of the plant cover of the National Park «Khvalynskiy». Scientifi c Works of National Park «Khvalynsky», iss. 1. Saratov, Khvalynsk, Nauchnaya kniga Publ., 2009, pp. 59–65 (in Russian).
4. Malysheva G. S., Malakhovskiy P. D. About Zonality of Vegetation on the Volga Upland (within the Saratov Region). Scientifi c Works of National Park «Khvalynsky», iss. 5. Saratov, Khvalynsk, ITs Nauka Publ., 2013, pp. 3–8 (in Russian).
5. Malakhovskiy P. D. Most interesting and in need of protection of the community of meadow steppes of the National Park «Khvalynsky». Scientifi c Works of National Park «Khvalynsky», iss. 2. Saratov, Izd-vo Sarat. tekhn. un-ta, 2010, pp. 49–57 (in Russian).
6. Malysheva G. S. Communities with Stipa lessingiana Trin. et Rupr. in the National Park «Khvalynsky» and the issues of their protection. Scientifi c Works of National Park «Khvalynsky», iss. 2. Saratov, Izd-vo Sarat. tekhn. un-ta, 2010, pp. 57–62 (in Russian).
7. Malysheva G. S., Malakhovskiy P. D., Suleymanova G. F. On the relationship between forest and steppe vegetation in the national park “Khvalynsky”. Scientifi c Works of National Park «Khvalynsky», iss. 3. Saratov, Khvalynsk, ITs Nauka Publ., 2011, pp. 30–43 (in Russian).
8. Braun-Blanquet J. Pfl anzensoziologie. Grundzuge der Vegetationskunde. 3. Aufl. Wien, New York, 1964. 865 S.
9. Chytry M., Hennekens S. M., Jimenez-Alfaro B., Knollova I., Dengler Ju., Jansen F., Landucci F., Schaminee J. H. J., Acic S., Agrillo E., Ambarli D., Angelini P., Apostolova I., Attore F., Berg C., Bergmeier E., Biurrun I., BottaDukat Z., Brisse H., Campos Ju. A., Carlon L., Carni A., Casella L., Csiky Ja., Custerevska R., Stephanovic Z. D., Danihelka J., De Bie E., de Ruffray P., De Sanctis M., Dickore W. B., Dimopoloulos P., Dubyna D., Dziuba T., Ejrn?s R., Ermakov N., Ewald J., Fanelli G., FernandezGonzalez F., FitzPatrick U., Font X., Garcia-Mijangos I., Gavilan R.G., Golub V., Guarino R., Haveman R., Indreica A., Gursoy D. I., Jandt U., Janssen J. A. M., Jirousek M., Kacki Z., Kavgaci A., Kleikamp M., Kolomiychuk V., Cuk M. K., Krstonosic D., Kuzemko A., Lenoir J., Lysenko T., Marceno K., Martynenko V., Michalkova D., Moeslund J. E., Onishchenko V., Pedashenko H., PerezHaase A., Peterka T., Prokhorov V., Rasomavicius V., Rodriguez-Rojo M. P., Rodwell J. S., Rogova T., Ruprecht E., Rusina S., Seidler G., Sibik J., Silc U., Skvorc Z., Sopotlieva D., Stancic Z., Svenning J.-Ch., Swacha G., Tsiripidis I., Turtureanu P.D., Ugurlu E., Uogintas D., Valachovic M., Vashenyak Yu., Vassilev K., Venanzoni R., Virtanen R., Weekes L., Willner W., Wohlgemuth Th., Yamalov S. European Vegetation Archive (EVA): an integrated database of European vegetation plots. Applied Vegetation Science, 2016, vol. 19, iss. 1, pp. 173–180. DOI: https://doi.org/10.1111/avsc.12191
10. Mucina L., Bultmann H., Dier?en K., Theurillat J.-P., Raus T., Carni A., Sumberova K., Willner W., Dengler J., Gavilan Garcia R., Chytry M., Hajek M., Di Pietro R., Iakushenko D., Pallas J., Daniels F. J. A., Bergmeier E., Santos Guerra A., Ermakov N., Valachovic M., Schaminee J. H. J., Lysenko T., Didukh Y. P., Pignatti S., Rodwell J. S., Capelo J., Weber H. E., Solomeshch A., Dimopoulos P., Aguiar C., Hennekens S. M., Tichy L. Vegetation of Europe: Hierarchical fl oristic classifi cation system of vascular plant, bryophyte, lichen, and algal communities. Applied Vegetation Science, 2016, vol. 19, iss. 1, pp. 3–264. DOI: https://doi.org/10.1111/avsc.12257
11. Lysenko T., Mitroshenkova A., Kalmykova O. Vegetation Database of the Volga and the Ural Rivers Basins. Vegetation Databases for the 21st Century. Biodiversity & Ecology, 2012, vol. 4, pp. 420–421. DOI: https://doi.org/10.7809.b-e.00208
12. Hennekens S. M. TURBO(VEG). Software package forinput, processing, and presentation of phytosociological data. Users guide. Version July 1996. IBN-DLO, Lancaster, 1996. 52 p.
13. Hill M. O. TWINSPAN – a FORTRAN program for arranging multivariate data in an ordered two-way table by classifi cation of the individuals and the attributes. Ithaca (NY), 1977. 48 p.
14. Tichy L. JUICE, software for vegetation classifi cation. J. Veg. Sci., vol. 13, pp. 451–453.
15. Mirkin B. M., Rozenberg G. S., Naumova L. G. Slovar’ ponyatiy i terminov sovremennoy fi totsenologii [The Dictionary of Notions and Terms in Modern Phytocenology]. Moscow, Nauka Publ., 1989. 223 p. (in Russian).
16. Cherepanov S. K. Sosudistyye rasteniya Rossii i sopredel’nykh gosudarstv (v predelakh byvshego SSSR) [Vascular plants of Russia and adjacent states (within the former USSR)]. St. Petersburg, Mir i semiya Publ., 1995. 992 p. (in Russian).
17. Weber H. E., Moravec J., Theurillat J.-P. International Code of Phytosociological Nomenclature. 3rd ed. J. Veg. Sci., 2000, vol. 11, pp. 739–768.
18. Klassifi katsiya i diagnostika pochv SSSR [Classifi cation and diagnostics of the soil of the USSR]. V. V. Egorov, E. N. Friedland, E. N. Ivanova, N. N. Rozov, V. A. Nosin, T. A. Friyev (compil.). Moscow, Kolos Publ., 1977. 224 p. (in Russian).
19. Grishin P. N., Kravchenko V. V., Boldyrev V. A. Pochvy Saratovskoy oblasti, ikh proiskhozhdeniye, sostav i agrokhimicheskiye svoystva [Soils of the Saratov region, Their Origin, Composition and Agrochemical Properties]. Saratov, Izd-vo Sarat. un-ta, 2011. 176 p. (in Russian).
20. Uzhametskaya E. A. Materialy k klassifi katsii lugovoy i stepnoy rastitel’nosti yuzhnoy chasti Samarskoy oblasti. II. Kharakteristika stepnoy rastitel’nosti (kl. FestucoBrometea) [Materials for the classifi cation of meadow and steppe vegetation in the southern part of the Samara area. II. Characteristics of steppe vegetation (cl. FestucoBrometea)], manuscript, deposited in 31.01.1992. 350- В92. Tol’yatti, 1992. 24 p. (in Russian).
21. Uzhametskaya E. A. Characteristics of the meadow and steppe vegetation of the valleys of the Bolshoy Irgiz and Bolshaya Glushitsa rivers (Samara Region). Bul. Samara Luka, 2002, no. 12, pp. 219–231 (in Russian).
22. Golub V. B., Saksonov S. V., Uzhametskaya E. A. Characterization of the stony steppes of the Zhiguli Mountains (based on the research of L. M. Cherepnin). Bul. Samara Luka, 1995, no. 6, pp. 73–96 (in Russian).
23. Golub V. B., Uzhametskaya E. A. Validization and brief characteristics of the seven syntaxons of the classes Molinio-Arrhenatheretea and Festuco-Brometea. Phytodiversity of Eastern Europe, 2016, vol. X, no. 2, pp. 197–205 (in Russian).
24. Lysenko T. M. To the characteristic of steppe vegetation of the Saratov area. Proceedings of the Samara Scientifi c Center of Russian Academy of Sciences, 2010, vol. 12, no. 1, pp. 61–66 (in Russian).
25. Lysenko T. M. Some Steppe Communities of the ForestSteppe Zone in the Volga Region. Problems of Steppe Science, 2014, no. XII, pp. 96–99 (in Russian).
26. Lysenko T. M. New Association of Steppe Vegetation from the Zhiguli Mountains. Bull. Samara Luka: Problems of Regional and Global Ecology, 2018, vol. 27, no. 1, pp. 213–217 (in Russian). DOI: https://doi.org/10.24411/2073-1035-2018-10012
27. Mitroshenkova A. E., Lysenko T. M. On the syntaxonomic characteristic of saucer-like karst craters in the Samara region. Local History Notes. Samara, SOIKM, 2004, vol. XIII, pp. 106–120 (in Russian).
28. Mitroshenkova A. E., Lysenko T. M. Syntaxonomic Characteristics of Plant Communities of Cone-shaped Plant Communities of Cone-Shaped Karst Reliefs in the Samara Region. Eastern Europe Phytodiversity, 2007, no. 4, pp. 26–52 (in Russian).
29. Mitroshenkova A. E., Lysenko T. M. New data on the vegetation cover of karst relief forms in the Samara region. News of the Samara Scientifi c Center of the Russian Academy of Sciences, 2009, vol. 11, no. 1 (4), pp. 638–642 (in Russian).
30. Demina O. N. Calcephyte Steppes of the Rostov Region. Vegetation of Russia, 2016, no. 29, pp. 21–45 (in Russian).
31. Petrova M. V., Lebedeva M. V., Yamalov S. M., Khasanova G. R. Syntaxonomy of the Pre-Ural Meadow Steppes. News of the Ufa Research Center of the Russian Academy of Sciences, 2018, no. 4, pp. 77–84 (in Russian). DOI: https://doi.org/10.31040/2222-8349-2018-0-4-77-84
32. Teptina A. YU., Lebedeva M. V., Yamalov S. M. On some communities of petrophytic steppes of the Middle Urals. Vegetation of Russia, 2018, no. 33, pp. 92–106 (in Russian). DOI: 10.31111/vegrus/2018.33.92
33. Poluyanov A. V., Averinova E. A. Travyanaya rasti tel’- nost’ Kurskoy oblasti (sintaksonomiya i voprosy okhrany) [Herbal Vegetation of the Kursk Region (syntaxonomy and protection issues)]. Kursk, Kursk State University, 2012. 276 p. (in Russian)